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Maize
Botanical illustration showing male and female flowers
Includes male and female flowers

Least Concern (IUCN 3.1)[2]
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Monocots
Clade: Commelinids
Order: Poales
Family: Poaceae
Subfamily: Panicoideae
Genus: Zea
Species:
Z. mays
Binomial name
Zea mays

Maize (

kernels or seeds, which are botanical fruits.[6][7] The term maize is preferred in formal, scientific, and international usage as the common name because this refers specifically to this one grain whereas corn refers to any principal cereal crop cultivated in a country. For example, in North America and Australia corn is often used for maize, but in England and Wales it can refer to wheat or barley, and in Scotland and Ireland to oats
.

Maize has become a staple food in many parts of the world, with the total production of maize surpassing that of wheat or rice. In addition to being consumed directly by humans (often in the form of masa), maize is also used for corn ethanol, animal feed and other maize products, such as corn starch and corn syrup.[8] The six major types of maize are dent corn, flint corn, pod corn, popcorn, flour corn, and sweet corn.[9] Sugar-rich varieties called sweet corn are usually grown for human consumption as kernels, while field corn varieties are used for animal feed, various corn-based human food uses (including grinding into cornmeal or masa, pressing into corn oil, fermentation and distillation into alcoholic beverages like bourbon whiskey), and as feedstocks for the chemical industry. Maize is also used in making ethanol and other biofuels.

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain.

Subsidies in the United States help to account for its high level of cultivation of maize and its position as the largest producer in the world.[12]

History

Pre-Columbian development

Cultivation of maize in an illustration from the 16th c. Florentine Codex
Ancient Mesoamerican relief, National Museum of Anthropology of Mexico

Maize is a

teosinte ancestor.[5] Genetic evidence can also be used to determine when various lineages split.[13]

Most historians believe maize was domesticated in the Tehuacán Valley of Mexico.[14] Recent research in the early 21st century has modified this view somewhat; scholars now indicate the adjacent Balsas River Valley of south-central Mexico as the center of domestication.[15]

An 2002 study by Matsuoka et al.. has demonstrated that, rather than the multiple independent domestications model, all maize arose from a single domestication in southern Mexico about 9,000 years ago. The study also demonstrated that the oldest surviving maize types are those of the Mexican highlands. Later, maize spread from this region over the Americas along two major paths. This is consistent with a model based on the archaeological record suggesting that maize diversified in the highlands of Mexico before spreading to the lowlands.[16][17]

Archaeologist Dolores Piperno has said:[15]

A large corpus of data indicates that [maize] was dispersed into lower Central America by 7600 BP [5600 BC] and had moved into the inter-Andean valleys of Colombia between 7000 and 6000 BP [5000–4000 BC].

— Dolores Piperno, The Origins of Plant Cultivation and Domestication in the New World Tropics: Patterns, Process, and New Developments[15]

Since then, even earlier dates have been published.[18]

According to a genetic study by the Brazilian Agricultural Research Corporation (Embrapa), corn cultivation was introduced in South America from Mexico, in two great waves: the first, more than 6000 years ago, spread through the Andes. Evidence of cultivation in Peru has been found dating to about 6700 years ago.[19] The second wave, about 2000 years ago, through the lowlands of South America.[20]

The earliest maize plants grew only small, 25-millimetre-long (1 in) corn ears, and only one per plant. In Jackson Spielvogel's view, many centuries of artificial selection (rather than the current view that maize was exploited by interplanting with

Olmec and Maya cultivated maize in numerous varieties throughout Mesoamerica; they cooked, ground and processed it through nixtamalization. It was believed that beginning about 2500 BC, the crop spread through much of the Americas.[22] Research of the 21st century has established even earlier dates. The region developed a trade network based on surplus and varieties of maize crops.[citation needed
]

potatoes in pre-Hispanic times; however, potato was the staple food of most Mapuches, "specially in the southern and coastal [Mapuche] territories where maize did not reach maturity".[23][24] Before the expansion of the Inca Empire maize was traded and transported as far south as 40°19' S in Melinquina, Lácar Department.[25] In that location maize remains were found inside pottery dated to 730 ± 80 BP and 920 ± 60 BP. Probably this maize was brought across the Andes from Chile.[25] The presence of maize in Guaitecas Archipelago (43°55' S), the southernmost outpost of pre-Hispanic agriculture,[26] is reported by early Spanish explorers.[27] However the Spanish may have misidentified the plant.[27]

By at least 1000 BCE, the

Olmec in Mesoamerica had based their calendar, language, myths and worldview with maize at the center of their symbolism.[28]

Columbian exchange

After the arrival of Europeans in 1492, Spanish settlers consumed maize, and explorers and traders

potatoes. Maize flour could not be substituted for wheat for communion bread, since in Christian belief only wheat could undergo transubstantiation and be transformed into the body of Christ.[29] Some Spaniards worried that by eating indigenous foods, which they did not consider nutritious, they would weaken and risk turning into Indians. "In the view of Europeans, it was the food they ate, even more than the environment in which they lived, that gave Amerindians and Spaniards both their distinctive physical characteristics and their characteristic personalities."[30] Despite these worries, Spaniards did consume maize. Archeological evidence from Florida sites indicate they cultivated it as well.[31]

Maize spread to the rest of the world because of its ability to grow in diverse climates. It was cultivated in Spain just a few decades after Columbus's voyages and then spread to Italy, West Africa and elsewhere.[31] Widespread cultivation most likely began in southern Spain in 1525, after which it quickly spread to the rest of the Spanish Empire including its territories in Italy (and, from there, to other Italian states). Maize had many advantages over wheat and barley; it yielded two and a half times the food energy per unit cultivated area,[32] could be harvested in successive years from the same plot of land, and grew in wildly varying altitudes and climates, from relatively dry regions with only 250 mm (10 in) of annual rainfall to damp regions with over 5,000 mm (200 in). By the 17th century it was a common peasant food in Southwestern Europe, including Portugal, Spain, southern France, and Italy. By the 18th century, it was the chief food of the southern French and Italian peasantry, especially in the form of polenta in Italy.[33]

Names

Many small male flowers make up the male inflorescence, called the tassel.

The word maize derives from the Spanish form of the indigenous

Taíno word for the plant, mahiz.[34] Linnaeus included the common name maize as the species epithet in Zea mays.[35] It is known by other names including "corn" in some English speaking countries.[36]

Maize is preferred in formal, scientific, and international usage as a common name because it refers specifically to this one grain, unlike corn, which has a complex variety of meanings that vary by context and geographic region.[37] The US and a handful of other English-speaking countries primarily use corn, though most countries use the term maize.[38][9][39] The word maize is considered interchangeable in place of corn in the West; during early British and American trade, all grains were considered corn. Maize retained the name corn in the West as the primary grain in these trade relationships.[35]

The word "corn" outside the US, Canada, Australia, and New Zealand is synonymous with grain referring to any cereal crop with its meaning understood to vary geographically to refer to the local staple,[40] such as wheat in England and oats in Scotland or Ireland.[37] In the United States,[40] Canada,[41] Australia, and New Zealand, corn primarily means maize. This usage started as a shortening of "Indian corn" in 18th century North America.[40][42] During European colonization of North America, confusion would occur between British and North American English speakers using the term corn so that North American speakers would need to clarify that they were talking about Indian corn or maize, such as in a conversation between the Massachusetts Bay governor Thomas Hutchinson and the British king George III.[42] "Indian corn" primarily means maize (the staple grain of indigenous Americans) but can also refer more specifically to multicolored "flint corn" used for decoration.[43] Other common names include barajovar, makka, silk maize, and zea.[44]

Betty Fussell writes in an article on the history of the word "corn" in North America that "[t]o say the word "corn" is to plunge into the tragi-farcical mistranslations of language and history".[28] Similar to the British, the Spanish referred to maize as panizo, a generic term for cereal grains, as did Italians with the term polenta. The British later referred to maize as Turkey wheat, Turkey corn, or Indian corn with Fusell commenting that "they meant not a place but a condition, a savage rather than a civilized grain", especially with Turkish people later naming it kukuruz, or barbaric.[28]

International groups such as the

FAO,[46] the International Maize and Wheat Improvement Center based out of Mexico, and the Indian Institute of Maize Research.[47] National agricultural and industry associations often include the word maize in their name such as the Maize Association of Australia,[48] and the National Maize Association of Nigeria.[49]

In Southern Africa, maize is commonly called mielie (Afrikaans) or mealie (English), words possibly derived from the Portuguese word for maize, milho, but more probably from Dutch meel or English meal, meaning the edible part of a grain or pulse.[50]

Structure and physiology

The maize plant is often 3 m (10 ft) in height,

internodes[54] of 18 cm (7 in) length.[51] The leaves arise from the nodes, alternately on opposite sides on the stalk,[55] and have entire margins.[56]

The apex of the stem ends in the tassel, an inflorescence of male flowers; these are separate from the female flowers but borne on the same plant (monoecy). When the tassel is mature and conditions are suitably warm and dry, anthers on the tassel dehisce and release pollen. Maize pollen is anemophilous (dispersed by wind), and because of its large settling velocity, most pollen falls within a few meters of the tassel.[57]

Ears develop above a few of the leaves in the midsection of the plant, between the stem and leaf sheath, elongating by around 3 mm (18 in) per day, to a length of 18 cm (7 in)[51] with 60 cm (24 in) being the maximum alleged in the subspecies.[58] They are female inflorescences, tightly enveloped by several layers of ear leaves commonly called husks.

Elongated

pericarp of the fruit is fused with the seed coat referred to as "caryopsis", typical of the grasses, and the entire kernel is often referred to as the "seed". The cob is close to a multiple fruit in structure, except that the individual fruits (the kernels) never fuse into a single mass. The grains are about the size of peas, and adhere in regular rows around a white, pithy substance, which forms the cob. The maximum size of kernels is reputedly 2.5 cm (1 in).[59] An ear commonly holds 600 kernels. They are of various colors: blackish, bluish-gray, purple, green, red, white and yellow. When ground into flour, maize yields more flour with much less bran than wheat does. It lacks the protein gluten of wheat and, therefore, makes baked goods with poor rising capability. A genetic variant that accumulates more sugar and less starch in the ear is consumed as a vegetable and is called sweet corn. Young ears can be consumed raw, with the cob and silk, but as the plant matures (usually during the summer months), the cob becomes tougher and the silk dries to inedibility. By the end of the growing season, the kernels dry out and become difficult to chew without cooking.[60]

  • Female inflorescence, with young silk
    Female inflorescence, with young silk
  • Mature silk
    Mature silk
  • Stalks, ears and silk
    Stalks, ears and silk
  • Male flowers
    Male flowers
  • Full-grown maize plants
    Full-grown maize plants
  • Mature maize ear on a stalk
    Mature maize ear on a stalk
Maize plant diagram

Planting density affects multiple aspects of maize. Modern farming techniques in developed countries usually rely on dense planting, which produces one ear per stalk.[61] Stands of silage maize are yet denser,[citation needed] and achieve a lower percentage of ears and more plant matter.[citation needed]

Maize is a

growing degree days > 10 °C (50 °F) in the environment to which it is adapted.[63] The magnitude of the influence that long nights have on the number of days that must pass before maize flowers is genetically prescribed[64] and regulated by the phytochrome system.[65] Photoperiodicity can be eccentric in tropical cultivars such that the long days characteristic of higher latitudes allow the plants to grow so tall that they do not have enough time to produce seed before being killed by frost. These attributes, however, may prove useful in using tropical maize for biofuels.[66]

Immature maize shoots accumulate a powerful antibiotic substance, 2,4-dihydroxy-7-methoxy-1,4-benzoxazin-3-one (

fungi. DIMBOA is also responsible for the relative resistance of immature maize to the European corn borer (family Crambidae). As maize matures, DIMBOA levels and resistance to the corn borer decline.[citation needed
]

Because of its shallow roots, maize is susceptible to droughts, intolerant of nutrient-deficient soils, and prone to be uprooted by severe winds.[67]

  • Maize kernels
    Maize kernels
  • Ear of maize with irregular rows of kernels
    Ear of maize with irregular rows of kernels

While yellow maizes derive their color from

suppressor.[73] The p1 gene encodes an Myb-homologous transcriptional activator of genes required for biosynthesis of red phlobaphene pigments, while the P1-wr allele specifies colorless kernel pericarp and red cobs, and unstable factor for orange1 (Ufo1) modifies P1-wr expression to confer pigmentation in kernel pericarp, as well as vegetative tissues, which normally do not accumulate significant amounts of phlobaphene pigments.[70] The maize P gene encodes a Myb homolog that recognizes the sequence CCT/AACC, in sharp contrast with the C/TAACGG bound by vertebrate Myb proteins.[74]

The ear leaf is the leaf most closely associated with a particular developing ear. This leaf and above contribute 70%[75] to 75% to 90%[76] of grain fill. Therefore fungicide application is most important in that region in most disease environments.[75][76]

  • cv. 'Ottofile giallo Tortonese'' – MHNT
    MHNT
  • cv. "strawberry"—MHNT
    cv. "strawberry"—
    MHNT
  • cv. "Oaxacan Green" MHNT
    cv. "Oaxacan Green"
    MHNT
  • Variegated maize ears
    Variegated maize ears
  • Multicolored corn kernels (CSIRO)
    Multicolored corn kernels (CSIRO)

Abnormal flowers

Maize flowers may sometimes exhibit mutations that lead to the formation of female flowers in the tassel. These mutations, ts4 and Ts6, prohibit the development of the stamen while simultaneously promoting pistil development.[77] This may cause inflorescences containing both male and female flowers, or hermaphrodite flowers.[78]

Genomics and genetics

selectively breeding new domestic strains
With white and yellow kernels

Maize is an annual grass in the family

perennial type of teosinte. The annual teosinte variety called Z. m. mexicana is the closest botanical relative to maize. It still grows in the wild as an annual in Mexico and Guatemala.[79]

Many forms of maize are used for food, sometimes classified as various subspecies related to the amount of starch each has:

This system has been replaced (though not entirely displaced) over the last 60 years by multivariable classifications based on ever more data.

Agronomic data were supplemented by botanical traits for a robust initial classification, then genetic, cytological, protein and DNA evidence was added. Now, the categories are forms (little used), races, racial complexes, and recently branches.[citation needed
]

Maize is a

transposon theory of "jumping genes", for which she won the 1983 Nobel Prize in Physiology or Medicine. Maize is still an important model organism for genetics and developmental biology today.[81]

The

kilobase. Kinetochores contain RNA originating from centromeres. Centromere regions can become inactive, and can continue in that state if the chromosome still has another active one.[82]

The Maize Genetics Cooperation Stock Center, funded by the USDA

tetraploids). Genetic data describing the maize mutant stocks as well as myriad other data about maize genetics can be accessed at MaizeGDB, the Maize Genetics and Genomics Database.[83]

In 2005, the US

B73 maize genome. The resulting DNA sequence data was deposited immediately into GenBank, a public repository for genome-sequence data. Sequences and genome annotations have also been made available throughout the project's lifetime at the project's official site.[84]

Primary sequencing of the maize genome was completed in 2008.

transposons, was found to contain 32,540 genes (By comparison, the human genome contains about 2.9 billion bases and 26,000 genes). Much of the maize genome has been duplicated and reshuffled by helitrons—group of rolling circle transposons.[87]

In Z. mays and various other angiosperms the MADS-box motif is involved in floral development. Early study in several angiosperm models including Z. mays was the beginning of research into the molecular evolution of floral structure in general, as well as their role in nonflowering plants.[88]

Varieties differ in their

CIMMYT maintains a large collection of maize/corn accessions tested and cataloged for insect resistance.[89]

single nucleotide polymorphism (SNP) typing for molecular breeding in this crop.[90] Semagn et al., 2013 describes methods and practices.[90]

Svitashev et al., 2016 demonstrates the integration of genome editing using (CRISPR–Cas9) into the breeding process for this crop.[91]

Evolution

As with many plants and animals, Z. mays has a positive correlation between

selection pressure. Z. m. having an EPS of ~650,000, it clusters with others of about the same EPS, and has 79% of its amino acid sites under selection.[92]

Recombination is a significant source of diversity in Z. mays. (Note that this finding supersedes previous studies which showed no such correlation.)[92]

This recombination/diversity effect is seen throughout plants but is also found to not occur – or not as strongly – in regions of high gene density. This is likely the reason that domesticated Z. mays has not seen as much of an increase in diversity within areas of higher density as in regions of lower density, although there is more evidence in other plants.[92]

Some lines of maize have undergone ancient

positive selection and lower negative selection which are observed in otherwise similar plants, i.e. also naturally outcrossing and with similar effective population sizes.[92]

Ploidy does not appear to influence EPS or magnitude of selection effect in maize.[92]

Breeding

Maize reproduces sexually each year. This randomly selects half the genes from a given plant to propagate to the next generation, meaning that desirable traits found in the crop (like high yield or good nutrition) can be lost in subsequent generations unless certain techniques are used.[citation needed]

Maize breeding in prehistory resulted in large plants producing large ears. Modern breeding began with individuals who selected highly productive varieties in their fields and then sold seed to other farmers. James L. Reid was one of the earliest and most successful developing Reid's Yellow Dent in the 1860s. These early efforts were based on

University of Illinois at Urbana-Champaign maintain germplasm important for future crop development.[citation needed
]

Since the 1940s the best strains of maize have been first-generation hybrids made from inbred strains that have been optimized for specific traits, such as yield, nutrition, drought, pest and disease tolerance. Both conventional cross-breeding and genetic engineering have succeeded in increasing output and reducing the need for cropland, pesticides, water and fertilizer.[94] There is conflicting evidence to support the hypothesis that maize yield potential has increased over the past few decades. This suggests that changes in yield potential are associated with leaf angle, lodging resistance, tolerance of high plant density, disease/pest tolerance, and other agronomic traits rather than increase of yield potential per individual plant.[95]

Certain varieties of maize have been bred to produce many ears which are the source of the "baby corn" used as a vegetable in Asian cuisine.[96]

One strain called olotón has evolved a symbiotic relationship with nitrogen-fixing microbes, which provides the plant with 29%–82% of its nitrogen.[97]

CIMMYT operates a conventional breeding program to provide optimized strains. The program began in the 1980s. Hybrid seeds are distributed in Africa by the Drought Tolerant Maize for Africa project.[94]

Genetic engineering

Bt maize. Grown since 1997 in the United States and Canada,[100] 92% of the US maize crop was genetically modified in 2016[98][101] and 33% of the worldwide maize crop was GM in 2016.[98][102] As of 2011, Herbicide-tolerant maize varieties were grown in Argentina, Australia, Brazil, Canada, China, Colombia, El Salvador, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, the Russian Federation, Singapore, South Africa, Taiwan, Thailand, and the United States. Insect-resistant maize was grown in Argentina, Australia, Brazil, Canada, Chile, China, Colombia, Egypt, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, South Africa, Switzerland, Taiwan, the United States, and Uruguay.[103]

In September 2000, up to $50 million worth of food products were recalled due to the presence of Starlink genetically modified corn, which had been approved only for animal consumption and had not been approved for human consumption, and was subsequently withdrawn from the market.[104]

For pest and disease resistance

Tropical

sorghum downy mildew resistance, and to further characterize SDMR regions already located by others.[106]

Origin

Teosinte
(top), maize-teosinte hybrid (middle), maize (bottom)

Maize is the

teosinte – teosintes are the crop wild relatives of this plant.[107] The two plants have dissimilar appearance, maize having a single tall stalk with multiple leaves and teosinte being a short, bushy plant. The difference between the two is largely controlled by differences in just two genes, called grassy tillers-1 (gt1, A0A317YEZ1) and teosinte branched-1 (tb1, Q93WI2).[107]

Several theories had been proposed about the specific origin of maize in Mesoamerica:[108][109]

  1. It is a direct domestication of a
    genetic material obtained from Z. m. ssp. mexicana through introgression.[16]
  2. It has been derived from
    hybridization between a small domesticated maize (a slightly changed form of a wild maize) and a teosinte of section Luxuriantes, either Z. luxurians or Z. diploperennis
    .
  3. It has undergone two or more domestications either of a wild maize or of a teosinte. (The term "teosinte" describes all species and subspecies in the genus Zea, excluding Z. m. ssp. mays.)
  4. It has evolved from a hybridization of Z. diploperennis by Tripsacum dactyloides.

In the late 1930s,

Paul Mangelsdorf suggested that domesticated maize was the result of a hybridization event between an unknown wild maize and a species of Tripsacum, a related genus. This theory about the origin of maize has been refuted by modern genetic testing, which refutes Mangelsdorf's model and the fourth listed above.[108]
: 40 

The teosinte origin theory was proposed by the Russian botanist

can cross-breed and produce fertile offspring
. A number of questions remain concerning the species, among them:

  1. how the immense diversity of the species of sect. Zea originated,
  2. how the tiny archaeological specimens of 3500–2700 BC could have been selected from a teosinte, and
  3. how domestication could have proceeded without leaving remains of teosinte or maize with teosintoid traits earlier than the earliest known until recently, dating from ca. 1100 BC.

The

archaeologists, geneticists, ethnobotanists, geographers, etc. The process is thought by some to have started 7,500 to 12,000 years ago. Research from the 1950s to 1970s originally focused on the hypothesis that maize domestication occurred in the highlands between the states of Oaxaca and Jalisco
, because the oldest archaeological remains of maize known at the time were found there.

Connection with 'parviglumis' subspecies

Genetic studies, published in 2004 by

Maya Maize God
, 550–850 AD

Doebley was part of the team that first published, in 2002, that maize had been domesticated only once, about 9,000 years ago, and then spread throughout the Americas.[16][115]

A primitive corn was being grown in southern Mexico, Central America, and northern South America 7,000 years ago. Archaeological remains of early maize ears, found at

Tehuacan, Puebla, 5,450 B.P.[22]

Maize pollen dated to 7,300 B.P. from San Andres, Tabasco, on the Caribbean coast has also been recovered.[113]

Jaina Island ceramic statuette of the young Maya Maize God emerging from an ear of corn, 600–900 A.D.

As maize was introduced to new cultures, new uses were developed and new varieties selected to better serve in those preparations. Maize was the staple food, or a major staple – along with

pre-Columbian North American, Mesoamerican, South American, and Caribbean cultures. The Mesoamerican civilization, in particular, was deeply interrelated with maize. Its traditions and rituals involved all aspects of maize cultivation – from the planting to the food preparation. Maize formed the Mesoamerican people's identity.[citation needed
]

It is unknown what precipitated its domestication, because the edible portion of the wild variety is too small, and hard to obtain, to be eaten directly, as each kernel is enclosed in a very hard bivalve shell.[citation needed]

In 1939, George Beadle demonstrated that the kernels of teosinte are readily "popped" for human consumption, like modern popcorn.[116] Some have argued it would have taken too many generations of selective breeding to produce large, compressed ears for efficient cultivation. However, studies of the hybrids readily made by intercrossing teosinte and modern maize suggest this objection is not well founded.[citation needed]

Spreading to the north

Around 4,500 years ago, maize began to spread to the north. Maize was first cultivated in what is now the United States at several sites in New Mexico and Arizona about 4,100 years ago.[117]

During the first millennium AD, maize cultivation spread more widely in the areas north. In particular, the large-scale adoption of maize agriculture and consumption in eastern North America took place about A.D. 900. Native Americans cleared large forest and grassland areas for the new crop.[118]

In 2005, research by the USDA Forest Service suggested that the rise in maize cultivation 500 to 1,000 years ago in what is now the southeastern United States corresponded with a decline of freshwater mussels, which are very sensitive to environmental changes.[119]

Cultivation

Planting

Seedlings three weeks after sowing
Young stalks

Because it is cold-intolerant, in the

soybeans. Maize is most sensitive to drought at the time of silk emergence, when the flowers are ready for pollination. In the United States, a good harvest was traditionally predicted if the maize was "knee-high by the Fourth of July", although modern hybrids generally exceed this growth rate. Maize used for silage is harvested while the plant is green and the fruit immature. Sweet corn is harvested in the "milk stage", after pollination but before starch has formed, between late summer and early to mid-autumn. Field maize is left in the field until very late in the autumn to thoroughly dry the grain, and may, in fact, sometimes not be harvested until winter or even early spring. The importance of sufficient soil moisture is shown in many parts of Africa, where periodic drought regularly causes maize crop failure and consequent famine. Although it is grown mainly in wet, hot climates, it has been said to thrive in cold, hot, dry or wet conditions, meaning that it is an extremely versatile crop.[120]

Mature plants showing ears

Maize was planted by the

squashes provided ground cover to stop weeds and inhibit evaporation by providing shade over the soil.[122] This method was replaced by single species hill planting where each hill 60–120 cm (2 ft 0 in – 3 ft 11 in) apart was planted with three or four seeds, a method still used by home gardeners. A later technique was "checked maize", where hills were placed 1 m (40 in) apart in each direction, allowing cultivators to run through the field in two directions. In more arid lands, this was altered and seeds were planted in the bottom of 10–12 cm (4–4+12 in) deep furrows to collect water. Modern technique plants maize in rows which allows for cultivation while the plant is young, although the hill technique is still used in the maize fields of some Native American reservations. When maize is planted in rows, it also allows for planting of other crops between these rows to make more efficient use of land space.[123]

In most regions today, maize grown in residential gardens is still often planted manually with a hoe, whereas maize grown commercially is no longer planted manually but rather is planted with a planter. In North America, fields are often planted in a two-crop rotation with a nitrogen-fixing crop, often alfalfa in cooler climates and soybeans in regions with longer summers. Sometimes a third crop, winter wheat, is added to the rotation.[citation needed]

Many of the maize varieties grown in the United States and Canada are hybrids. Often the varieties have been

genetically modified to tolerate glyphosate or to provide protection against natural pests. Glyphosate is an herbicide which kills all plants except those with genetic tolerance. This genetic tolerance is very rarely found in nature.[citation needed
]

In the midwestern United States, low-till or

fertilized. Weeds are controlled through the use of herbicides
, and no cultivation tillage is done during the growing season. This technique reduces moisture evaporation from the soil, and thus provides more moisture for the crop. The technologies mentioned in the previous paragraph enable low-till and no-till farming. Weeds compete with the crop for moisture and nutrients, making them undesirable.[citation needed]

Harvesting

Mature maize ears
Harvesting maize, Jones County, Iowa
Rantasalmi, South Savonia
, Finland
Hand-picking harvest of maize in Myanmar

Maize harvested as a grain crop can be kept in the field a relatively long time, even months, after the crop is ready to harvest; it is also harvested and stored in the husk leaves if kept dry.[124] Before the 20th century, all maize harvesting was by manual labour, by grazing, or by some combination of those. Whether the ears were hand-picked and the stover was grazed, or the whole plant was cut, gathered, and shocked, people and livestock did all the work. Between the 1890s and the 1970s, the technology of maize harvesting expanded greatly. Today, all such technologies, from entirely manual harvesting to entirely mechanized, are still in use to some degree, as appropriate to each farm's needs, although the thoroughly mechanized versions predominate, as they offer the lowest unit costs when scaled to large farm operations.

Before World War II, most maize in North America was harvested by hand. This involved a large number of workers and associated social events (husking or shucking bees). From the 1890s onward, some machinery became available to partially mechanize the processes, such as one- and two-row mechanical pickers (picking the ear, leaving the stover) and corn binders, which are reaper-binders designed specifically for maize. The latter produce sheaves that can be shocked. By hand or mechanical picker, the entire ear is harvested, which then requires a separate operation of a maize sheller to remove the kernels from the ear. Whole ears of maize were often stored in corn cribs, and these whole ears are a sufficient form for some livestock feeding use. Today corn cribs with whole ears, and corn binders, are less common because most modern farms harvest the grain from the field with a combine and store it in bins. The combine with a corn head (with points and snap rolls instead of a reel) does not cut the stalk; it simply pulls the stalk down. The stalk continues downward and is crumpled into a mangled pile on the ground, where it usually is left to become organic matter for the soil. The ear of maize is too large to pass between slots in a plate as the snap rolls pull the stalk away, leaving only the ear and husk to enter the machinery. The combine separates the husk and the cob, keeping only the kernels.[125]

The entire maize plant is also harvested as a silage crop.[126]

For storing grain in bins, the moisture of the grain must be sufficiently low to avoid spoiling. If the moisture content of the harvested grain is too high, grain dryers are used to reduce the moisture content by blowing heated air through the grain. This can require large amounts of energy in the form of combustible gases (propane or natural gas) and electricity to power the blowers.[127]

Production

Production of maize (2019)[128]

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain.[10] In 2020, total world production was 1.16 billion tonnes, led by the United States with 31.0% of the total (table). China produced 22.4% of the global total.[129]

Top Maize producers
in 2020
Numbers in million
FAOSTAT[130][129]

United States

In 2016, maize production was forecast to be over 380 million metric tons (15 billion

Midwestern states such as Indiana, Iowa, and Illinois; in the latter, it was named the state's official grain in 2017.[132]

The estimated corn usage for crop year September 1, 2020 to August 31, 2021, was 38.7 percent was used for feed, 34 percent for ethanol, 17.5 percent for export, and 9.8 percent for food.[133]

Trade

Corn futures are traded on several exchanges, the Chicago Board of Trade (CBOT) and JSE Derivatives (JDERIV). The Chicago Board Of Trade sells corn futures with a contact size of 5000 bushels which is quoted in cents/bushel and the JDERIV has a contact size of 100 Tonnes, quoted in Rand/Ton.[134][135] The detailed contract specifications are listed below:

Contract Specifications[134][135]
Corn (CORN) Corn (CNA)
Exchange: JDERIV CBOT
Sector: Grain Grain
Tick Size: 0.2 0.25
Tick Value: 5 ZAR 12.50 USD
BPV: 25 50
Denomination: ZAR USD
Decimal Place: 1 2

Pests

Insects

The susceptibility of maize to the European corn borer and corn rootworms, and the resulting large crop losses which are estimated at a billion dollars worldwide for each pest,

transgenics expressing the Bacillus thuringiensis
toxin. "Bt maize" is widely grown in the United States and has been approved for release in Europe.

Diseases

Storage

Drying is vital to prevent or at least reduce mycotoxin contamination. Aspergillus and Fusarium spp. are the most common mycotoxin sources, but there are others. Altogether maize contaminants are so common, and this crop is so economically important, that maize mycotoxins are among the most important in agriculture in general.[100]

Uses

Culinary

Semi-peeled corn on the cob
Poster showing a woman serving muffins, pancakes, and grits, with canisters on the table labeled corn meal, grits, and hominy, US Food Administration, 1918

Maize and

grain alcohol. Grain alcohol from maize is traditionally the source of Bourbon whiskey. Cornmeal and maize flour are used to make cornbread and other baked products. Many countries require corn flour to be enriched with certain nutrients
.

In prehistoric times

]

Maize is a staple of

huitlacoche, is considered a delicacy.[citation needed
]

tamales
made with corn meal
Boiled corn on a white plate

Coarse maize meal is made into a thick

mieliepap in South Africa and sadza, nshima, ugali and other names in other parts of Africa. Introduced into Africa by the Portuguese in the 16th century, maize has become Africa's most important staple food crop.[144] These are commonly eaten in the Southeastern United States, foods handed down from Native Americans, who called the dish sagamite.[citation needed
]

Maize can also be harvested and consumed in the unripe state, when the kernels are fully grown but still soft. Unripe maize must usually be cooked to become palatable; this may be done by simply boiling or roasting the whole ears and eating the kernels right off the cob. Sweet corn, a genetic variety that is high in sugars and low in starch, is usually consumed in the unripe state. Such corn on the cob is a common dish in the United States, Canada, United Kingdom, Cyprus, some parts of South America, and the Balkans, but virtually unheard of in some European countries.[citation needed] Corn on the cob was hawked on the streets of early 19th-century New York City by poor, barefoot "Hot Corn Girls", who were thus the precursors of hot dog carts, churro wagons, and fruit stands seen on the streets of big cities today.[145]

Within the United States, the usage of maize for human consumption constitutes only around 1/40th of the amount grown in the country. In the United States and Canada, maize is mostly grown to feed livestock, as forage, silage (made by fermentation of chopped green cornstalks), or grain. Maize meal is also a significant ingredient of some commercial animal food products.[citation needed]

Nutritional value

Sweetcorn, yellow, raw
(seeds only)
Note: assuming freed niacin
Nutritional value per 100 g (3.5 oz)
Energy360 kJ (86 kcal)
18.7 g
Starch5.7 g
Sugars6.26 g
Dietary fiber2 g
1.35 g
3.27 g
Tryptophan0.023 g
Threonine0.129 g
Isoleucine0.129 g
Leucine0.348 g
Lysine0.137 g
Methionine0.067 g
Cystine0.026 g
Phenylalanine0.150 g
Tyrosine0.123 g
Valine0.185 g
Arginine0.131 g
Histidine0.089 g
Alanine0.295 g
Aspartic acid0.244 g
Glutamic acid0.636 g
Glycine0.127 g
Proline0.292 g
Serine0.153 g
Niacin (B3)
11%
1.77 mg
Pantothenic acid (B5)
14%
0.717 mg
Vitamin B6
5%
0.093 mg
Folate (B9)
11%
42 μg
Vitamin C
8%
6.8 mg
MineralsQuantity
%DV
Iron
3%
0.52 mg
Magnesium
9%
37 mg
Manganese
7%
0.163 mg
Phosphorus
7%
89 mg
Potassium
9%
270 mg
Zinc
4%
0.46 mg
Other constituentsQuantity
Water75.96 g

Link to USDA Database entry
One ear of medium size (6-3/4" to 7-1/2" long)
maize has 90 grams of seeds
Percentages estimated using US recommendations for adults,[146] except for potassium, which is estimated based on expert recommendation from the National Academies.[147]

Raw, yellow, sweet maize kernels are composed of 76% water, 19%

essential minerals, magnesium and phosphorus whereas other nutrients are in low amounts (table).[citation needed
]

Maize has suboptimal amounts of the essential amino acids tryptophan and lysine, which accounts for its lower status as a protein source.[148] However, the proteins of beans and legumes complement those of maize.[148]

Feed and fodder for livestock

Maize is a major source of both grain feed and fodder for livestock. It is fed to the livestock in various ways. When it is used as a grain crop, the dried kernels are used as feed. They are often kept on the cob for storage in a corn crib, or they may be shelled off for storage in a grain bin. The farm that consumes the feed may produce it, purchase it on the market, or some of both. When the grain is used for feed, the rest of the plant (the corn stover) can be used later as fodder, bedding (litter), or soil amendment. When the whole maize plant (grain plus stalks and leaves) is used for fodder, it is usually chopped all at once and ensilaged, as digestibility and palatability are higher in the ensilaged form than in the dried form. Maize silage is one of the most valuable forages for ruminants.[149] Before the advent of widespread ensilaging, it was traditional to gather the corn into shocks after harvesting, where it dried further. With or without a subsequent move to the cover of a barn, it was then stored for weeks to several months until fed to the livestock. Today ensilaging can occur not only in siloes but also in silage wrappers. However, in the tropics, maize can be harvested year-round and fed as green forage to the animals.[150]

Chemicals

Starch from maize can also be made into

fabrics, adhesives, and many other chemical products.[citation needed
]

The

wet milling process, is widely used in the biochemical industry and research as a culture medium to grow many kinds of microorganisms.[151]

Chrysanthemin is found in purple corn and is used as a food coloring.[citation needed]

Bio-fuel

"Feed maize" is being used increasingly for heating;[152] specialized corn stoves (similar to wood stoves) are available and use either feed maize or wood pellets to generate heat. Maize cobs are also used as a biomass fuel source. Maize is relatively cheap and home-heating furnaces have been developed which use maize kernels as a fuel. They feature a large hopper that feeds the uniformly sized maize kernels (or wood pellets or cherry pits) into the fire.[citation needed]

Maize is increasingly used as a feedstock for the production of ethanol fuel.[153] When considering where to construct an ethanol plant, one of the site selection criteria is to ensure there is locally available feedstock.[154] Ethanol is mixed with gasoline to decrease the amount of pollutants emitted when used to fuel motor vehicles. High fuel prices in mid-2007 led to higher demand for ethanol, which in turn led to higher prices paid to farmers for maize. This led to the 2007 harvest being one of the most profitable maize crops in modern history for farmers. Because of the relationship between fuel and maize, prices paid for the crop now tend to track the price of oil. [citation needed]

The price of food is affected to a certain degree by the use of maize for biofuel production. The cost of transportation, production, and marketing are a large portion (80%) of the price of food in the United States. Higher energy costs affect these costs, especially transportation. The increase in food prices the consumer has been seeing is mainly due to the higher energy cost. The effect of biofuel production on other food crop prices is indirect. Use of maize for biofuel production increases the demand, and therefore price of maize. This, in turn, results in farm acreage being diverted from other food crops to maize production. This reduces the supply of the other food crops and increases their prices.[155][156]

Farm-based maize silage digester located near Neumünster in Germany, 2007. The green tarpaulin top cover is held up by the biogas stored in the digester.

Maize is widely used in Germany as a feedstock for

biogas plants. Here the maize is harvested, shredded then placed in silage clamps from which it is fed into the biogas plants. This process makes use of the whole plant rather than simply using the kernels as in the production of fuel ethanol.[citation needed
]

A

Increasingly, ethanol is being used at low concentrations (10% or less) as an additive in

biofuels" and has been generating an intense debate regarding the human beings' necessity of new sources of energy, on the one hand, and the need to maintain, in regions such as Latin America, the food habits and culture which has been the essence of civilizations such as the one originated in Mesoamerica; the entry, January 2008, of maize among the commercial agreements of NAFTA has increased this debate, considering the bad labor conditions of workers in the fields, and mainly the fact that NAFTA "opened the doors to the import of maize from the United States, where the farmers who grow it receive multimillion-dollar subsidies and other government supports. ... According to OXFAM UK, after NAFTA went into effect, the price of maize in Mexico fell 70% between 1994 and 2001. The number of farm jobs dropped as well: from 8.1 million in 1993 to 6.8 million in 2002. Many of those who found themselves without work were small-scale maize growers.").[158] However, introduction in the northern latitudes of the US of tropical maize for biofuels, and not for human or animal consumption, may potentially alleviate this.[citation needed
]

Commodity

Maize is bought and sold by investors and price speculators as a tradable commodity using corn futures contracts. These "futures" are traded on the Chicago Board of Trade (CBOT) under ticker symbol C. They are delivered every year in March, May, July, September, and December.[159]

Ornamental and other uses

Some forms of the plant are occasionally grown for ornamental use in the garden. For this purpose, variegated and colored leaf forms as well as those with colorful ears are used.[citation needed]

Corncobs can be hollowed out and treated to make inexpensive

smoking pipes, first manufactured in the United States in 1869.[citation needed
]

Children playing in a maize kernel box

An unusual use for maize is to create a "

hedges, but these take several years to mature. The rapid growth of a field of maize allows a maze to be laid out using GPS at the start of a growing season and for the maize to grow tall enough to obstruct a visitor's line of sight by the start of the summer. In Canada and the US, these are popular in many farming communities.[citation needed
]

Maize kernels can be used in place of sand in a sandboxlike enclosure for children's play.[161]

Stigmas from female maize flowers, popularly called

herbal supplements.[citation needed
]

Maize is used as a fish bait, called "dough balls". It is particularly popular in Europe for coarse fishing.[citation needed]

Additionally, feed corn is sometimes used by hunters to bait animals such as deer or wild hogs.[citation needed]

United States usage breakdown

The breakdown of usage of the 12.1-billion-bushel (307-million-tonne) 2008 US maize crop was as follows, according to the World Agricultural Supply and Demand Estimates Report by the USDA.[162]

Use Amount
million bushels million tonnes percentage
livestock feed 5,250 133.4 43.4
ethanol production 3,650 92.7 30.2
exports 1,850 47.0 15.3
production of starch, corn oil, sweeteners (HFCS, etc.) 943 24.0 7.8
human consumption—grits, corn flour, corn meal, beverage alcohol 327 8.3 2.7

In the US since 2009/2010, maize feedstock use for ethanol production has somewhat exceeded direct use for livestock feed; maize use for fuel ethanol was 5,130 million bushels (130 million tonnes) in the 2013/2014 marketing year.[163]

A fraction of the maize feedstock dry matter used for ethanol production is usefully recovered as DDGS (dried distillers grains with solubles). In the 2010/2011 marketing year, about 29.1 million tonnes of DDGS were fed to US livestock and poultry.[164] Because starch utilization in fermentation for ethanol production leaves other grain constituents more concentrated in the residue, the feed value per kg of DDGS, with regard to ruminant-metabolizable energy and protein, exceeds that of the grain. Feed value for monogastric animals, such as swine and poultry, is somewhat lower than for ruminants.[164]

Comparison to other staple foods

The following table shows the nutrient content of maize and major staple foods in a raw harvested form on a dry weight basis to account for their different water contents. Raw forms are not usually eaten and cannot be digested well. They are either sprouted, or prepared and cooked for human consumption. In sprouted or cooked form, the nutritional and anti-nutritional contents of each of these staples differ from that of raw form of these staples reported in the table below.

Note: niacin for maize assumes freed niacin.
Nutrient content of 10 major staple foods per 100 g dry weight[169]
Staple Maize (corn)[A] Rice, white[B] Wheat[C] Potatoes[D] Cassava[E] Soybeans, green[F] Sweet potatoes[G] Yams[Y] Sorghum[H] Plantain[Z]
RDA
Water content (%) 10 12 13 79 60 68 77 70 9 65
Raw grams per 100 g dry weight 111 114 115 476 250 313 435 333 110 286
Nutrient
Energy (kJ) 1698 1736 1574 1533 1675 1922 1565 1647 1559 1460 8,368–10,460
Protein (g) 10.4 8.1 14.5 9.5 3.5 40.6 7.0 5.0 12.4 3.7 50
Fat (g) 5.3 0.8 1.8 0.4 0.7 21.6 0.2 0.6 3.6 1.1 44–77
Carbohydrates (g) 82 91 82 81 95 34 87 93 82 91 130
Fiber (g) 8.1 1.5 14.0 10.5 4.5 13.1 13.0 13.7 6.9 6.6 30
Sugar (g) 0.7 0.1 0.5 3.7 4.3 0.0 18.2 1.7 0.0 42.9 minimal
Minerals [A] [B] [C] [D] [E] [F] [G] [Y] [H] [Z]
RDA
Calcium (mg) 8 32 33 57 40 616 130 57 31 9 1,000
Iron (mg) 3.01 0.91 3.67 3.71 0.68 11.09 2.65 1.80 4.84 1.71 8
Magnesium (mg) 141 28 145 110 53 203 109 70 0 106 400
Phosphorus
(mg)
233 131 331 271 68 606 204 183 315 97 700
Potassium (mg) 319 131 417 2005 678 1938 1465 2720 385 1426 4700
Sodium (mg) 39 6 2 29 35 47 239 30 7 11 1,500
Zinc
(mg)
2.46 1.24 3.05 1.38 0.85 3.09 1.30 0.80 0.00 0.40 11
Copper
(mg)
0.34 0.25 0.49 0.52 0.25 0.41 0.65 0.60 - 0.23 0.9
Manganese
(mg)
0.54 1.24 4.59 0.71 0.95 1.72 1.13 1.33 - - 2.3
Selenium (μg) 17.2 17.2 81.3 1.4 1.8 4.7 2.6 2.3 0.0 4.3 55
Vitamins [A] [B] [C] [D] [E] [F] [G] [Y] [H] [Z]
RDA
Vitamin C (mg) 0.0 0.0 0.0 93.8 51.5 90.6 10.4 57.0 0.0 52.6 90
Thiamin
(B1) (mg)
0.43 0.08 0.34 0.38 0.23 1.38 0.35 0.37 0.26 0.14 1.2
Riboflavin (B2) (mg) 0.22 0.06 0.14 0.14 0.13 0.56 0.26 0.10 0.15 0.14 1.3
Niacin
(B3) (mg)
4.03 1.82 6.28 5.00 2.13 5.16 2.43 1.83 3.22 1.97 16
Pantothenic acid (B5) (mg) 0.47 1.15 1.09 1.43 0.28 0.47 3.48 1.03 - 0.74 5
Vitamin B6 (mg) 0.69 0.18 0.34 1.43 0.23 0.22 0.91 0.97 - 0.86 1.3
Folate Total (B9) (μg) 21 9 44 76 68 516 48 77 0 63 400
Vitamin A (IU) 238 0 10 10 33 563 4178 460 0 3220 5000
Vitamin E, alpha-tocopherol (mg) 0.54 0.13 1.16 0.05 0.48 0.00 1.13 1.30 0.00 0.40 15
Vitamin K1
(μg)
0.3 0.1 2.2 9.0 4.8 0.0 7.8 8.7 0.0 2.0 120
Beta-carotene
(μg)
108 0 6 5 20 0 36996 277 0 1306 10500
Lutein+zeaxanthin (μg) 1506 0 253 38 0 0 0 0 0 86 6000
Fats [A] [B] [C] [D] [E] [F] [G] [Y] [H] [Z]
RDA
Saturated fatty acids (g) 0.74 0.20 0.30 0.14 0.18 2.47 0.09 0.13 0.51 0.40 minimal
Monounsaturated fatty acids (g) 1.39 0.24 0.23 0.00 0.20 4.00 0.00 0.03 1.09 0.09 22–55
Polyunsaturated fatty acids (g) 2.40 0.20 0.72 0.19 0.13 10.00 0.04 0.27 1.51 0.20 13–19
[A] [B] [C] [D] [E] [F] [G] [Y] [H] [Z]
RDA

A raw yellow dent corn
B raw unenriched long-grain white rice
C raw hard red winter wheat
D raw potato with flesh and skin
E raw cassava
F raw green soybeans
G raw sweet potato
H raw sorghum
Y raw yam
Z raw plantains
/* unofficial

Hazards

Pellagra

When maize was first introduced into farming systems other than those used by traditional native-American peoples, it was generally welcomed with enthusiasm for its productivity. However, a widespread problem of malnutrition soon arose wherever maize was introduced as a staple food. This was a mystery, since these types of malnutrition were not normally seen among the indigenous Americans, for whom maize was the principal staple food.[170]

It was eventually discovered that the indigenous Americans had learned to soak maize in

niacin, the lack of which was the underlying cause of the condition known as pellagra.[171]

Maize was introduced into the diet of non-indigenous Americans without the necessary cultural knowledge historically acquired in the Americas. In the late 19th century, pellagra reached epidemic proportions in parts of the southern US, as medical researchers debated two theories for its origin: the deficiency theory (which was eventually shown to be true) said that pellagra was due to a deficiency of some nutrient, and the germ theory said that pellagra was caused by a germ transmitted by stable flies. Another theory promoted by the eugenicist Charles Davenport held that people only contracted pellagra if they were susceptible to it due to certain "constitutional, inheritable" traits of the affected individual.[172]

Once alkali processing and dietary variety were understood and applied, pellagra disappeared in the developed world. The development of high lysine maize and the promotion of a more balanced diet have also contributed to its demise. Pellagra still exists today in food-poor areas and refugee camps where people survive on donated maize.[173]

Allergy

Maize contains

mucous membranes, diarrhea, vomiting, asthma and, in severe cases, anaphylaxis. It is unclear how common this allergy is in the general population.[citation needed
]

The Z. mays plant has an

OPALS allergy scale rating of 5 out of 10, indicating moderate potential to cause allergic reactions, exacerbated by over-use of the same plant throughout a garden. Corn pollen is heavy, large, and usually airborne in the early morning.[175]

Mycotoxins

Bt maize discourages insect vectors and by so doing it dramatically reduces concentrations of fumonisins, significantly reduces aflatoxins, but only mildly reduces others.[100]

Art

Gold maize. Moche culture 300 A.D., Larco Museum, Lima, Peru
Water tower in Rochester, Minnesota being painted as an ear of maize

Maize has been an essential crop in the Andes since the pre-Columbian era. The Moche culture from Northern Peru made ceramics from earth, water, and fire. This pottery was a sacred substance, formed in significant shapes and used to represent important themes. Maize was represented anthropomorphically as well as naturally.[176]

In the United States, maize ears along with tobacco leaves are carved into the capitals of columns in the United States Capitol building. Maize itself is sometimes used for temporary architectural detailing when the intent is to celebrate the fall season, local agricultural productivity and culture. Bundles of dried maize stalks are often displayed along with pumpkins, gourds and straw in autumnal displays outside homes and businesses. A well-known example of architectural use is the Corn Palace in Mitchell, South Dakota, which uses cobs and ears of colored maize to implement a mural design that is recycled annually. Another well-known example is the Field of Corn sculpture in Dublin, Ohio, where hundreds of concrete ears of corn stand in a grassy field.[177]

A maize stalk with two ripe ears is depicted on the reverse of the Croatian 1 lipa coin, minted since 1993.[178]

Shucked, a 2022 musical that is currently running on Broadway, was described by Vulture as a "show about corn". Much of the show contains puns relating to corn[179][180] and the plot revolves around a blighted corn crop.[181]

See also

References

  1. ^ https://en.wikipedia.org/wiki/Maize
  2. ^ Contreras, A., Ruíz Corral, J.A., Menjívar, J., Aragón Cuevas, F., González Ledesma, M. & Sánchez, J.J. 2019. Zea mays. The IUCN Red List of Threatened Species 2019: e.T77726273A77726310. https://dx.doi.org/10.2305/IUCN.UK.2019-2.RLTS.T77726273A77726310.en. Accessed on 28 October 2022.
  3. ^ "maíz". Diccionario de la lengua española - Edición del Tricentenario (in Spanish). Retrieved January 14, 2023.
  4. ^ "The Evolution of Corn". University of Utah HEALTH SCIENCES. Retrieved January 2, 2016.
  5. ^
    PMID 11172083
    .
  6. ^ "Please settle a dispute. Is sweet corn a vegetable or a grain? What is the difference? How about field corn? - eXtension". USDA National Institute of Food and Agriculture, New Technologies for Ag Extension project. Archived from the original on June 21, 2018. Retrieved March 3, 2018.
  7. ^ Chodosh, Sara (July 8, 2021). "The bizarre botany that makes corn a fruit, a grain, and also (kind of) a vegetable". Popular Science. Retrieved February 24, 2022.
  8. ^ Foley, Jonathon. "It's Time to Rethink America's Corn System". Scientific American. Retrieved February 18, 2019.
  9. ^ . Retrieved February 15, 2023. The use of the word "corn" for what is termed "maize" by most other countries is peculiar to the United States. Europeans who were accustomed to the names "wheat corn," "barley corn," and "rye corn" for other small-seeded cereal grains referred to the unique American grain maize as "Indian corn." The term was shortened to just "corn," which has become the American word for the plant of American genesis.
  10. ^ (PDF).
  11. ^ Genetically modified plants: Global Cultivation Area Maize Archived August 12, 2010, at the Wayback Machine GMO Compass, March 29, 2010, retrieved August 10, 2010
  12. ^ "Corn Production By State 2021". worldpopulationreview.com. Retrieved July 16, 2021.
  13. .
  14. ^ "Origin, History and Uses of Corn". Iowa State University, Department of Agronomy. February 11, 2014. Archived from the original on February 23, 2014.
  15. ^
    S2CID 83061925
    . Recent studies in the Central Balsas River Valley of Mexico, maize's postulated cradle of origin, document the presence of maize phytoliths and starch grains at 8700 BP, the earliest date recorded for the crop (Piperno et al. 2009; Ranere et al. 2009). A large corpus of data indicates that it was dispersed into lower Central America by 7600 BP and had moved into the inter-Andean valleys of Colombia between 7000 and 6000 BP. Given the number of Cauca Valley, Colombia, sites that demonstrate early maize, it is likely that the inter-Andean valleys were a major dispersal route for the crop after it entered South America
  16. ^ .
  17. ^ Matsuoka, Yoshihiro (January 22, 2003). "Earliest Directional Evolution for Microsatellite Size in Maize" (PDF). Science. Retrieved March 3, 2014.
  18. .
  19. ^ "Los antiguos peruanos comían palomitas de maíz". BBC Mundo. BBC. January 19, 2012.
  20. ^ "Did man follow plants or did plants follow man? Tracks of prehistoric man and ways of contact in the Americas according to cultivated plants. Case study – Maize (translated from Portuguese)". Yumpu. 2015. Retrieved October 13, 2015.
  21. .
  22. ^ a b Roney, John (Winter 2009). "The Beginnings of Maize Agriculture". Archaeology Southwest. 23 (1): 4.
  23. .
  24. .
  25. ^ .
  26. ^ Bird, Junius (1946). "The Alacaluf". In Steward, Julian H. (ed.). Handbook of South American Indians. Bulletin 143. Vol. I. –Bureau of American Ethnology. pp. 55–79.
  27. ^ .
  28. ^ . To say the word "corn" is to plunge into the tragi-farcical mistranslations of language and history. If only the British had followed Columbus in phoneticizing the Taino word mahiz, which the Arawaks named their staple grain, we wouldn't be in the same linguistic pickle we're in today, where I have to explain to someone every year that when Biblical Ruth "stood in tears amid the alien corn" she was standing in a wheat field. But it was a near thing even with the Spaniards, when we read in Columbus' Journals that the grain "which the Indians called maiz... the Spanish called panizo.' The Spanish term was generic for the cereal grains they knew - wheat, millet, barley, oats - as was the Italian term polenta, from Latin pub. As was the English term "corn," which covered grains of all kinds, including grains of salt, as in "corned beef.
    French linguistic imperialism, by way of a Parisian botanist in 1536, provided the term Turcicum frumentum, which the British quickly translated into "Turkey wheat," "Turkey corn," and "Indian corn." By Turkey or Indian, they meant not a place but a condition, a savage rather than a civilized grain, with which the Turks concurred, calling it kukuruz, meaning barbaric.
  29. ^ Rebecca Earle, The Body of the Conquistador: Food, Race, and the Colonial Experience in Spanish America, 1492–1700. New York: Cambridge University Press 2012, pp. 17, 151.
  30. ^ Earle, The Body of the Conquistador, p. 5.
  31. ^ a b Earle, The Body of the Conquistador, p. 144.
  32. ^ Marion Eugene Ensminger and Audrey H. Ensminger. "Foods & Nutrition Encyclopedia, Two Volume Set." CRC-Press: 1994. Page 1104.
  33. ^ William L. Langer, "American Foods and Europe's Population Growth 1750–1850", Journal of Social History, 8#2 (1975), pp. 51–66. Pages 58–60.
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    S2CID 4640742
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  36. .
  37. ^ . The word "maize" is preferred in international usage because in many countries the term "corn", the name by which the plant is known in the United States, is synonymous with the leading cereal grain; thus, in England "corn" refers to wheat, and in Scotland and Ireland it refers to oats.
  38. ^ Espinoza, Mauricio. "'All Corn Is the Same,' and Other Foolishness about America's King of Crops". Ohio State University: College of Food, Agricultural, and Environmental Sciences. Retrieved September 21, 2022.
  39. ^ "Grain". education.nationalgeographic.org. National Geographic. Retrieved February 27, 2023. In most countries, the grain of the Zea mays plant is called maize. In the United States, it's called corn.
  40. ^ a b c "corn, n.1". Oxford English Dictionary (Online ed.). Oxford University Press. (Subscription or participating institution membership required.)
  41. .
  42. ^ . Corn, in orthodox English, means grain for human consumption, especially wheat, e.g., the Corn Laws. The earliest settlers, following this usage, gave the name of Indian corn to what the Spaniards, following the Indians themselves, had called maiz. . . . But gradually the adjective fell off, and by the middle of the Eighteenth Century maize was simply called corn and grains in general were called breadstuffs. Thomas Hutchinson, discoursing to George III in 1774, used corn in this restricted sense speaking of "rye and corn mixed." "What corn?" asked George. "Indian corn," explained Hutchinson, "or as it is called in authors, maize."
  43. ^ "Indian corn", Merriam-Webster Dictionary, definition 3, accessed June 7, 2012
  44. S2CID 87844060
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  45. ^ "Zea mays (maize)". CABI. Retrieved September 16, 2022.
  46. ^ "Maize". FAO. Retrieved September 16, 2022.
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Further reading

External links


Category:Zea (plant) Category:Agriculture in Mesoamerica Category:Crops originating from Mexico Category:Demulcents Category:Energy crops Category:Flora of Guatemala Category:Flora of Mexico Category:Fruit vegetables Category:Grasses of Mexico Category:Plant models Category:Pre-Columbian Native American cuisine Category:Post-Columbian Native American cuisine Category:Pre-Columbian Southwest cuisine Category:Staple foods Category:Tropical agriculture Category:Plants described in 1753 Category:Symbols of Illinois